Two years later, a molecular analysis showed that " E . gracilis " was, in fact, more closely related to " Astasia longa " than to certain other species recognized as " Euglena ".
2.
If any doubt remained, it was dispelled in 1994, when genetic analysis of the non-photosynthesizing euglenoid " Astasia longa " confirmed that this organism retains sequences of DNA inherited from an ancestor that must have had functioning chloroplasts.