As is usual for the genus, the molars are brachyodont, low-crowned, and bunodont, with the cusps higher than the connecting crests, and accessory crests such as mesolophs are present.
12.
The molars are bunodont ( with the cusps higher than the connecting crests ) and brachyodont ( low-crowned ) and have the inter-cusp valleys on the labial ( outer ) sides closed by a cingulum ( shelf ).