Diagnostic features are : uniformity, rather than asymmetry, of the rings of the ocelli; an undivided, dorsally domed uncus in the male genitalia; a short broad aedeagus with two apical fields of scobination, and a densely and robustly scobinate apex to the conical vesica; strong pouches lateral to the ostium in the female genitalia; a signum consisting of a single spine in the bursa copulatrix.