According to Suh et al . ( 2011 Science 334 : 1415 ) the projection to CA3 and dentate gyrus in mice is primarily from layer II of entorhinal cortex, and forms a trisynaptic path with hippocampus ( dentate gyrus to CA3 to CA1 ), distinguished from the direct ( monosynaptic ) perforant path from Layer III of entorhinal cortex to CA1 and subiculum.