First species of this genus, " Osperleioceras bicarinatum " has evolved in what is now France from " Harpoceras subplanatum ".
2.
In 1884 he received his doctorate in natural sciences from the University of Strasbourg with a dissertation on the ammonite genus " Harpoceras ", titled " " Beitr�ge zu einer monographie der Ammonitengattung Harpoceras " ".
3.
In 1884 he received his doctorate in natural sciences from the University of Strasbourg with a dissertation on the ammonite genus " Harpoceras ", titled " " Beitr�ge zu einer monographie der Ammonitengattung Harpoceras " ".
4.
They were collected from the " Harpoceras semicelatum-elegantulum-exaratum " ammonoid subzones ( Lias ? II1-5, about 182.7-181.8 mya ), " Harpoceras tenuicostatum-falcifer " zones, of the Posidonia Shale.
5.
They were collected from the " Harpoceras semicelatum-elegantulum-exaratum " ammonoid subzones ( Lias ? II1-5, about 182.7-181.8 mya ), " Harpoceras tenuicostatum-falcifer " zones, of the Posidonia Shale.