Using karyotypes, Defler has done work clarifying the taxonomy of various species of night monkey ( " Colombian Amazon on the white-fronted capuchin ( " Cebus albifrons " ), the brown woolly monkey ( " Lagothrix lagothricha " ), the black-headed uakari ( " Cacajao melanocephalus " ), the black titi ( " Callicebus lugens " ), the Lucifer titi ( " Callicebus lucifer " ), and the Venezuelan red howler ( " Alouatta seniculus " ) and has accomplished many primate censuses in different parts of eastern Colombia.
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Using karyotypes, Defler has done work clarifying the taxonomy of various species of night monkey ( " Colombian Amazon on the white-fronted capuchin ( " Cebus albifrons " ), the brown woolly monkey ( " Lagothrix lagothricha " ), the black-headed uakari ( " Cacajao melanocephalus " ), the black titi ( " Callicebus lugens " ), the Lucifer titi ( " Callicebus lucifer " ), and the Venezuelan red howler ( " Alouatta seniculus " ) and has accomplished many primate censuses in different parts of eastern Colombia.