Stein's scheme was amended in 1889 by Otto B�tschli, who divided the Ciliata into two orders : Holotricha and Spirotricha.
2.
In his system, Spirotricha is retained as an order, under the class Polyhymenophora, but Holotricha does not appear at any rank.
3.
The other three orders of Stein's Ciliata were placed in a lower rank under the new order Spirotricha : ciliates possessing a well-developed structure of spiralling membranelles around the oral cavity.
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